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University of Nevada, Reno
University of
Nevada, Reno

Dr. C. Richard Tracy, Professor
Autecology, Desert Biology, Paleoecology, and Conservation Biology

California State University, Northridge
1966, B.A., Biology
1968, M.S., Biology


University of Wisconsin
1972, Ph.D., Major Zoology, Minor Botany

University of Nevada
Biology Department m/s 314
Reno NV 89557

email Dr. Richard Tracy

Office phone
775-784-1925
FAX number
775-784-1302

Current Projects / Research Interests:

My research interests and experiences are broad and include descriptive ecology, autecology (broadly defined to include physiological ecology, biophysical ecology, animal behavior, and population biology), evolutionary biology, theoretical ecology, organismal biology, and conservation biology. Some of my research interests include: Proximate and ultimate interactions between organisms and their environments. The ecological importance of Grinnellian Niches in organisms, and the autecological function of these niche relationships in determining distribution, dispersion, dispersal, and population fluctuations. Conservation Biology: the ecology and conservation of amphibians and reptiles (especially sensitive species such as desert tortoise and western toads); the biology, ecology, and conservation of deserts; and paleobiology and extinction process.

Selected Publications:

Tracy C.R
., K. E. Nussear, T. C. Esque, K. Dean-Bradley, C. R. Tracy, L. A. DeFalco, K. T. Castle, L. C. Zimmerman, R. E. Espinoza, and A. M. Barber. 2006. The importance of physiological ecology in conservation biology. Integrative and Comparative Biology. Integrative and Comparative Biology, pp. 1-15. doi:10.1093/icb/icl054. This is an Open Access article (http://creativecommons.org/licenses/by-nc/2.0/uk/) which permits unrestricted non-commercial use, distribution, and reproduction in any medium, provided the original work is properly cited.

Tracy, C.R
., Christian, K.A., Reynolds, S.J., McArthur, L., Tracy, C.R. 2006 Ecology of aestivation in an Australian frog, Cyclorana australis (Hylidae). Copeia (in press).

Christian, K. A., Tracy, C.R., Tracy, C.R. 2006. Evaluating thermoregulation in reptiles: an appropriate null model. The American Naturalist 168(3): electronic journal article.

Nussear, K.E. and C. R. Tracy. 2006. Can modeling of desert tortoise activity improve estimation of population densities. Ecological Applications 17(2): 579-586.

Espinoza, R.E., J.J. Wiens, and C.R. Tracy. 2004. Novel ecophysiological adaptations promote multiple origins of herbivory in a group of South American lizards. Proceedings of the National Academy of Science. 101(48): 16819-16824

Simandle, E.T., R.E. Espinoza, K.E. Nussear, and C.R. Tracy. 2001. Lizards, lipids, and dietary links to animal function. Physiological and Biochemical Zoology 74(5): 625-640.

Tartowski, S.L., E.B. Allen, N.E. Barrett, A.R. Berkowitz, R.K. Colwell, P.M. Groffman, J. Harte, H.P. Possingham, C.M. Pringle, D.L. Strayer, and C.R. Tracy. 1996. Toward a resolution of conflicting paradigms. In: The Ecological Basis of Conservation: Heterogeneity, Ecosystems, and Biodiversity, S.T.A. Pickett, R.S. Ostfield, M. Shachak, and G.E. Likens (Eds.), Chapman & Hall, NY.

Tracy, C.R. and T.L. George. 1992. On the determinants of extinction. American Naturalist 139(1): 102 122.

Tracy, C.R., J.S. Turner, and R.B. Huey. 1986. A biophysical analysis of possible thermoregulatory adaptations in pelycosaurs. Pp. 195-205 In: Ecology and Biology of Mammal-Like Reptiles, P.D. MacLean, J.J. Roth, E.C. Roth, and N. Hotton (Eds.), Smithsonian Press, Washington, DC.

Tracy, C.R. 1982. Biophysical modelling in reptilian physiology and ecology. In: Biology of the Reptilia, C. Gans and F.H. Pough (Eds.), Academic Press, London, Volume 12, pp 275-321.

Tracy, C.R. 1977. Minimum size of mammalian homeotherms: The role of the thermal environment. Science 198: 1034-1035.

Tracy, C.R. 1976. A model of the dynamic exchanges of water and energy between a terrestrial amphibian and its environment. Ecological Monographs 46(3): 293-326.

Additional Links:

Biodiversity at UNR
Ecology, Evolution, and Conservation Biology Program

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